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SRX4277302: GSM3208739: WT_p53-ChIP; Danio rerio; ChIP-Seq
1 ILLUMINA (Illumina HiSeq 4000) run: 30.9M spots, 2.3G bases, 819.9Mb downloads

Submitted by: NCBI (GEO)
Study: P53 based ChIP-sequencing of wildtype and mutant supt16h-/- zebrafish
show Abstracthide Abstract
Hematopoietic stem cells (HSCs) have long been the focus of developmental and regenerative studies, yet our understanding of the signaling events regulating their specification remains incomplete. We demonstrate that supt16h, a component of the FAcilitates Chromatin Transcription (FACT) complex, is required for HSC formation. Zebrafish supt16h mutants express reduced levels of Notch signaling components, genes essential for HSC development, due to abrogated transcription. Classically, Supt16h regulates transcription and nucleosome reorganization. Whereas global chromatin accessibility in supt16h mutants is unaffected, we observe a specific increase in accessibility at the p53 locus, causing an accumulation of p53 mRNA and protein. We further demonstrate that P53 levels directly influence expression of the Polycomb Group protein, phc1, which functions as a transcriptional repressor of Notch genes. Suppression of phc1 or its upstream regulator, p53, rescues both loss of Notch and loss of HSC phenotypes in supt16h mutants. Taken together, our results highlight a previously uncharacterized relationship between supt16h, p53, and phc1 to specify HSCs via modulation of Notch signaling. Overall design: ChIP-seq for the P53 protein was performed on roughly 1,000 of each wildtype (WT) and supt16h-/- whole embryos at 32hpf.
Sample: WT_p53-ChIP
SAMN09462749 • SRS3443390 • All experiments • All runs
Organism: Danio rerio
Library:
Instrument: Illumina HiSeq 4000
Strategy: ChIP-Seq
Source: GENOMIC
Selection: ChIP
Layout: SINGLE
Construction protocol: Approximately 1,000 of each WT and MUT supt16h embryos per treatment were collected and deyolked using the Ginzburg Fish Ringer solution. Embryos were resuspended in 1 mL PBS with 1 mM PMSF, 1X cOmplete EDTA Free Protease Inhibitor Cocktail Tablet (Roche), 1X phosSTOP Phosphatase Inhibitor Tablet (Roche) and pushed through a 21G needle. Embryos were immediately crosslinked with 4 mM DSG (ProteoChem) for 30 min, followed by 1% PFA for 8 min, and stopped with 0.125M Glycine. Fixed embryos were resuspended in 1 mL Lysis Buffer 1 (0.05M HEPES, 140 mM NaCl, 1 mM EDTA, 1% Glycerol, 0.5% NP40, 0.25% Triton-X) and subjected to the TissueRuptorII (Qiagen) for 20 sec on low. Tissue was rocked for 10 min at 4ºC, pelleted, and resuspended in 1 mL Lysis Buffer 2 (10 mM Tris,-pH 8, 200 mM NaCl, 1mM EDTA, 0.5 mM EGTA). They were rocked again for 10 min at room temperature (RT), pelleted and resuspended in 1.5 mL Lysis Buffer 3 (10mM Tris-pH 8, 100 mM NaCl, 1 mM EDTA, 0.5 mM EGTA, 0.1% sodium deoxycholate, 0.05% SDS). Tissue was then sonicated on the Diagenode Bioruptor 300 (30 sec ON, 30 sec OFF, HIGH, 34 cycles). Following the addition of 0.8% Triton-X to the sonicated chromatin, cells were spun down. 20 ul of lysate was saved as input control and the remainder was incubated with Dyna Beads bound with 10 ug p53 antibody (GeneTex) overnight at 4ºC, rotating. Beads were collected on a magnet and washed 1x Wash Buffer 1 (20 mM Tris-pH 8, 150 mM NaCl, 2 mM EDTA. 0.1% SDS, 1% Triton-X), 1x Wash Buffer 2 (20mM Tris-pH 8, 500 mM NaCl, 2 mM EDTA, 0.1% SDS, 1% Triton-X), 1x Wash Buffer 3 (10mM Tris-pH 8, 250 mM LiCl, 1 mM EDTA, 1% sodium deoxycholate, 1% NP40), 3x TET Buffer (1x TE, 0.2% Tween-20), and 1x TE-NaCl (1x TE, 50 mM NaCl). Beads were then resuspended in 500 ul Elution Buffer (50mM NaHCO3, 1% SDS) and incubated overnight with 0.2M NaCl at 65ºC to reverse crosslinking. Lysate was then incubated for 2 hrs at 37ºC with 5 ul RNase A (10 mg/ml) and 2 hrs at 55ºC with 5 ul Proteinase K (20 mg/ml). DNA was purified using Phenol/Chroloform/Isoamyl and resuspended in 50 ul TE (10 mM Tris-pH 8, 1 mM EDTA). TruSeq ChIP Library Preparation Kit (Illumina) was used to construct libraries which were subsequently run at SR75 on the HiSeq4000.
Experiment attributes:
GEO Accession: GSM3208739
Links:
Runs: 1 run, 30.9M spots, 2.3G bases, 819.9Mb
Run# of Spots# of BasesSizePublished
SRR740522730,907,7612.3G819.9Mb2020-01-02

ID:
5764687

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